FAMILIA GEOTRUPIDAE Latreille, 1802
SUBFAMILIA BOLBOCERATINAE Mulsant, 1842
Tribu Athyreini Lynch Arribálzaga, 1878 / Howden & Martinez, 1963
Neoathyreus (Neoathyreus) excavatus (Laporte de Castelnau, 1840)
Neoathyreus (Neoathyreus) fissicornis (Harold, 1880) [= Athyreus fissicornis Harold, 1880]
Neoathyreus (Neoathyreus) granulicollis Howden, 1964
Neoathyreus (Neoathyreus) interruptus Howden, 1964
Neoathyreus (Neoathyreus) mexicanus (Klug, 1845) [= Athyreus mexicanus Klug, 1845]
Neoathyreus (Neoathyreus) mixtus Howden, 1964
Tribu Bolboceratini Mulsant, 1842
Bolbelasmus arcuatus (Bates, 1887:111) [= Bolboceras arcuatus Bates, 1887]
Bolbelasmus bajaensis Howden, 1964:44
Bolbelasmus carinifrons Howden & Solis, 1995:261
Bolbelasmus horni (Rivers, 1886) [= Bradycinetus horni Rivers, 1886]
Bolbelasmus rotundipennis
Bolbelasmus variabilis
Bolbocerastes imperialis imperialis Cartwright, 1953:109
Bolbocerastes peninsularis (Schaeffer, 1906) [= Bradycinetus peninsularis Schaeffer, 1906]
Bolbocerastes regalis Cartwright, 1953:106
Bolbocerastes serratus (LeConte, 1854) [= Athyreus serratus LeConte, 1854]
Bolbocerosoma mexicanum Howden, 2005
Bolbocerosoma pusillum townesi Howden, 1955:183
Bolbocerosoma ritcheri Howden, 1955:186
Bolborhombus magnus Howden, 1964
Bolborhombus nitidus Howden, 1964
Bolborhombus parvulus Cartwright, 1953
Bolborhombus sallaei (Bates, 1887) [= Bolboceras sallaei Bates, 1887; = Bradycinctus carinatus Schaeffer, 1906; = Bolboceras schaefferi Boucomont, 1911]
Eucanthus greeni Robinson, 1948:30
Eucanthus impressus Howden, 1964:72
Eucanthus mexicanus Howden, 1964
SUBFAMILIA GEOTRUPINAE Latreille, 1802
Tribu Geotrupini Latreille, 1802
Ceratotrupes bolivari Halffter & Martinez, 1962
Ceratotrupes fronticornis (Erichson, 1847) [= Geotrupes fronticornis Erichson, 1847]
Ceratotrupes sturmi Jekel, 1866 [= Geotrupes mniszechi Jekel, 1866]
Cnemotrupes cnephosus Howden, 1964
Cnemotrupes guerreroensis Howden, 1964 [in subgenus Onthotrupes]
Cnemotrupes herbeus Jekel, 1866 [in subgenus Onthotrupes]
Cnemotrupes lobatus Howden, 1974 [in subgenus Onthotrupes]
Cnemotrupes nebularum Howden, 1964 [ in subgenus Onthotrupes]
Cnemotrupes pecki Howden, 1974 [in subgenus Onthotrupes]
Cnemotrupes sallaei (Jekel, 1866) [= Geotrupes sallaei Jekel, 1866] [= Geotrupes falsus Jekel, 1866] [as variety of sallei]
Cnemotrupes sobrinus Jekel, 1866 [in subgenus Onthotrupes] [= Geotrupes sobrina Jekel, 1866]
Cnemotrupes truncaticornis Howden, 1964
Cnemotrupes viridiobscurus Jekel, 1866 [in subgenus Onthotrupes] [= Geotrupes felschei Nonfried, 1894] [= Geotrupes saundersi Jekel, 1866]
Halffterius rufoclavatus (Jekel, 1866) [= Geotrupes rufoclavatus Jekel, 1866]
Haplogeotrupes guatemalensis unidentatus Howden, 1974
Haplogeotrupes reddelli (Howden, 1980) [= Cnemotrupes reddelli Howden, 1980]
Megatrupes cavicollis (Bates, 1887) [= Geotrupes cavicollis Bates, 1887]
Megatrupes fisheri (Howden, 1967) [= Geotrupes fisheri Howden, 1967]
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BIBLIOGRAFÍA
Blackwelder, R.E., 1944. (Taurocerastinae, Geotrupinae) Checklist of the coleopterous insects of México, Central America, The West Indies, and South America. U.S. National Museum Bull., 185 (2):219-220
Boilly, Olivier. (Geotrupidae, Bolboceratidae). Listado de publicaciones.
Bouchard, Patrice; Yves Bousquet; Anthony E. Davies; Miguel A. Alonso-Zarazaga; John F. Lawrence; Chris H. C. Lyal; Alfred F. Newton; Chris A. M. Reid; Michael Schmitt; S. Adam Ślipiński; Andrew B. T. Smith. 2011. Family-group names in Coleoptera (Insecta).ZooKeys 88: 1–972 (april, 2011)
Lawrence, J. F. and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 in: J. Pakaluk and S.A. Slipinski (eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa.
Leschen, R. A. B.; Beutel, R. G. and Lawrence, J. F. (Editors); Slipinski, A. (Associate editor). 2009. Handbook of zoology. Coleoptera, Beetles, vol. 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).
TAMU (Texas A & M University). http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Geotrupidae.txt
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